Neutral and acid detergent fibres were measured in order to determine the insoluble fractions. Estimates of lignin, neutral detergent fibre (NDF), and acid detergent fibre (ADF) were made using a slightly modified version of the Van Soest (1967) and McQueen and Nicholson (1979) methods. The following formulas were used to estimate the contents of hemicellulose and cellulose: hemicellulose = NDF-ADF and cellulose = ADF-lignin, respectively.