Morphological terminology for the body parts of the beetles is following the work of Leschen [2 ].
Photographs were taken at the Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine (Kiev, SIZK) using the microscope Leica Z16 APO stereomicroscope equipped with a Leica DFC 450 camera. Dimensions were measured by 10 × 23 Leica 10450023Widefield Adjustable Eyepiece on Leica MZ APO.
Eleven extant specimens cited in the text were collected by late Alexey Yu. Isaev (Ulyanovsk State University, Russia) and are deposited in the Zoological Museum of Lomonosov State University (Moscow, ZMSU).
The holotype is deposited in collection of Schmalhausen Institute of Zoology.