Previous studies on pumas have explored behavior time budgets [13] , minimum area requirements [14] , dispersal of juveniles in fragmented habitat [15] , and the relationship between urban development and puma presence [16] , [17] . We expand upon these studies by using GPS data combined with extensive field reconnaissance to determine the spatial location of pumas while exhibiting four behaviors chosen to reflect both reproductive and non-reproductive activities: feeding, moving, communicating, and denning, and how these are influenced by natural and anthropogenic factors.
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